Ecological Archives A022-014-A1

Andrea S. Downing, Egbert H. van Nes, Jan H. Janse, Frans Witte, Ilse J. M. Cornelissen, Marten Scheffer, and Wolf M. Mooij. 2012. Collapse and reorganization of a food web of Mwanza Gulf, Lake Victoria. Eological Applications 22:229–239.

Appendix A. A description of the input parameters needed to create an ECOPATH mass balance, their units, and empirical relations necessary to calculate them, following the ECOPATH user guide (Christensen et al. 2005).

TABLE A1. Input parameters needed to create an ECOPATH mass-balance.

Parameter Description Unit
Biomass (B) Average biomass per unit area in the habitat area where the group occurs t/km2
Production to
biomass ratio (P / B)
Equivalent to the instantaneous rate of total mortality (Z) y-1
Consumption to
biomass (Q / B)
Rate of intake of food by a group over time y-1
Ecotrophic
efficiency (EE)
Fraction of production that is further used in the system  
Production over
consumption (P / Q)
Ratio between production and consumption,
corresponds to gross food conversion efficiency
 
Unassimilated
consumption
Fecal fraction, part of consumption that returns to detritus  
Detritus import Rate of entry of detritus into the system t/km2/y
Fishing landings Represent the non-discarded part of the catch, expressed as flows t/km2/y
Diets Expressed as the fraction each prey contributes to the overall diet of a group  

TABLE A2. Empirical equations used to calculate productivity and consumption (Christensen et al. 2005), as well as quantifying Caridina nilotica* stocks (Goudswaard et al. 2006).

Productivity P / B = Z = M + F
P / B = K × (L - L) / (L - L')
Consumption Q / B = 10C1 - W-C2 × 10-C3 × 10A × C4
Natural mortality M = KC5 × L-C6 × C7
Fishing mortality F = Y / B
Length L = Lt / (1 - e(t - t0))
Weight-length W = a × Lb
Shrimp weight-length
relationship
*
FW = LC8 × e-C9 / C10

TABLE A3. Parameters needed for the empirical equations.

P Production t/km2/y
B Biomass t/km2
Z Total mortality rate y-1
M Natural mortality rate y-1
F Fishing mortality rate y-1
K von Bertalanffy growth function parameter expressing
rate at which asymptotic length is approached
y-1
L8 Asymptotic length cm
L Mean length cm
L’ Length at entrance into fishery cm
Y Yield t/km2
Q Consumption t/km2/y
W8 Asymptotic weight G
A Tail aspect ratio parameter  
Lt Length at age t Cm
t Age y
t0 von Bertalanffy growth function parameter
expressing hypothetical age of a fish of length 0
y
W Weight g
L Length cm
a Scaling factor in W-L relationship  
b Exponent in W-L relationship  
DW Dry weight G
FW Fresh weight g

TABLE A4. Constants used in empirical relations (Christensen et al. 2005).

C1= 7.964 C6= 0.279
C2= 0.204 C7= 4.439
C3= 6.583 C8= 2.7
C4= 0.083 C9= 4.876
C5= 0.65 C10=0.2
 

LITERATURE CITED

Christensen, V., C. Walters, and D. Pauly. 2005. ECOPATH with ECOSIM: a user's guide. Fisheries Centre, University of British Columbia, Vancouver.

Goudswaard, K. P. C., F. Witte, and J. H. Wanink. 2006. The shrimp Caridina nilotica in Lake Victoria (East africa), before and after the Nile perch increase. Hydrobiologia 563:31–44.


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