Ecological Archives E088-008-A2

Alan R. Townsend, Cory C. Cleveland, Gregory P. Asner, and Mercedes M. C. Bustamante. 2007. Controls over foliar N:P ratios in tropical rain forests. Ecology 88:107–118.

Appendix B. Mean foliar N:P values, latitude, and mean annual rainfall for 45 evergreen tropical forests. In some cases, multiple sites are reported for the same general area (e.g., San Carlos, Venezuela, Maui, Panama, the Osa Peninsula of Costa Rica); each of these represent distinct sites that differ from each other in some significant state factor (e.g., rainfall and/or soil type).

Country/State

Mean annual
rainfall (mm)

Absolute latitude
(°N or S)

Foliar N:P

Reference

Zaire

1700

0.5

20.4

Bartholomew et al. 1953

Brazil

2500

1.0

21.9

Johnson et al. 2001

Malaysia

4200

1.4

43.5

Peace and Macdonald 1981

Venezuela

3565

1.9

21.2

Medina 1984

Venezuela

3565

1.9

29.7

Klinge 1984

Venezuela

3565

2.0

16.6

Medina 1984

Venezuela

3565

2.0

18.0

Klinge 1984

Venezuela

3565

2.0

14.8

Medina 1984

Venezuela

3565

2.0

11.4

Cuevas and Klinge 1986

Venezuela

3565

2.0

10.8

Cuevas and Klinge 1986

Venezuela

3565

2.0

22.3

Cuevas and Klinge 1986

Brazil

3600

2.0

22.2

Klinge 1985

Ecuador

3000

2.2

19.1

Davidson et al. 1999

Brazil

2100

2.4

36.8

Klinge 1984

Brazil

2200

2.5

29.3

This paper

Brazil

1600

3.2

16.9

Thompson et al. 1992

Brazil

2100

3.4

30.0

This paper

Colombia

1988

4.0

10.8

Vera et al. 1999

Colombia

2400

4.0

7.5

Vera et al. 1999

Cameroon

5060

5.0

17.0

Chuyong et al. 2000

Ghana

1500

6.1

18.0

Greenland and Kowal 1960; Nye 1958

Colombia

3000

6.4

27.6

Folster et al. 1976

Venezuela

1700

7.3

16.9

Hase and Folster 1982

Venezuela

1500

8.4

15.2

Grimm and Fassbender 1981

Costa Rica

5200

8.4

16.4

This paper

Costa Rica

5200

8.4

13.3

This paper

Brazil

2500

8.5

25.4

Martinelli et al. 2000

Panama

1800

9.0

16.4

Santiago et al. 2005

Panama

2300

9.0

21.7

Santiago et al. 2005

Panama

3500

9.0

21.5

Santiago et al. 2005

Panama

3100

9.0

18.7

Santiago et al. 2005

New Guinea

4000

10.0

16.5

Grubb and Edwards 1982

Costa Rica

4000

10.3

12.8

Hiremath et al. 2002

Malawi

1400

16.0

11.8

Maghembe and Prins 1994

Jamaica

2500

18.0

17.9

Tanner 1977

China

1700

18.4

24.3

Han et al. 2005

Puerto Rico

3500

18.4

18.5

Sanchez et al. 1997

Puerto Rico

3500

18.4

23.0

Sanchez et al. 1997

Hawaii

2500

19.0

18.5

Vitousek et al. 1995

China

1900

19.1

9.5

Han et al. 2005

Maui

2200

19.8

20.2

Schuur and Matson 2001

Maui

2450

19.8

23.0

Schuur and Matson 2001

Maui

2750

19.8

18.4

Schuur and Matson 2001

Maui

3350

19.8

19.5

Schuur and Matson 2001

Maui

4050

19.8

17.5

Schuur and Matson 2001

Maui

5050

19.8

11.5

Schuur and Matson 2001

Kauai

2500

21.0

16.2

Vitousek et al. 1995

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